(Sep 2007)  Page(s) 345-346.  
 
Rosa nutkana C. Presl. subsp. melina (Greene) W. H. Lewis & Ertter, comb. et stat. nov. Basionym: Rosa melina Greene, Pittonia 4: 10. 1899. TYPE: USA. Colorado: Montrose Co., Cerro Summit, near Cimarron, 30 Aug 1896. E. L. Greene s.n. (holotype, NDG 23707). Figure 1E-H.
....Rosa nutkana subsp. melina is endemic to high mountains of Colorado, New Mexico, and Utah and rare in southeastern Idaho, Montana, and Wyoming, often on isolated peaks and ridges in or at edges of aspen, fir, spruce, and/or pine forest at elevations of 2500-3800+ m. This distribution, to some degree, includes the subspecies in the South Rocky Mountain Floristic Element...., paralleling that of R. woodsii subsp. manca except for the Mogollon Rim component. In some regions, as in central and western Colorado and southeastern Utah, R. nutkana subsp. melina is frequent and may be found growing near or with R. woodsii subsp. manca, but the subspecies also occurs at higher elevations beyond the others' range. Hybriods are possible but not expected even in mixed or nearby populations; this is because R. nutkana subsp. melina is hexaploid, while R. woodsii subsp. manca is diploid and because flowering of R. nutkana subsp. melina occurs before that of R. woodsii subsp. manca, although there can be an overlap of anthesis. No intermediate tetraploid hybrid has been found.

(Sep 2007)  Page(s) 345.  
 
Rosa nutkana C. Presl. subsp. melina (Greene) W. H. Lewis & Ertter, comb. et stat. nov. ....
Rosa oreophila Rydberg. Bull. Torrey Bot. Club 31: 561. 1904. Syn. nov. [new Synonym] TYPE: USA. Colorado: Routt Co., Four-miles Hill, 2590 m. 22 July 1896. C. F. Baker s.n. (holotype, NY 415860; isotypes, MO 1951203, NDC 23560, NY 415862).

(Sep 2007)  Page(s) 345-346.  
 
Rosa nutkana C. Presl. subsp. melina (Greene) W. H. Lewis & Ertter, comb. et stat. nov. ...
Rosa underwoodii Rydberg, Bull. Torrey Bot. Club 31: 560. 1904. Syn. nov. [new synonym]TYPE: USA. Colorado: Ouray Co., W. of Ouray, above Box añon. 2300-2700 m. 8 Sep. 1904. L. M. Underwood & A. D. Selby 122 (holotype, NY 429915).

(10 Sep 2007)  Page(s) 346-347.  Includes photo(s).
 
Rosa woodsii Lindley, Ros. Monogr. 21. 1820....
Key to the subspecies of Rosa woodsii in Central and Western North America:
- Shrubs most frequently short, typically less than 1 m, infrequently to 2 m; fertile branches densely prickly of various sizes, rarely few or unarmed; terminal leaflets obovate, fewer ovate or elliptic; flowers 1 to 3, rarely more; prairies and plains of central North America extending into the Rocky Mtns. and nearby Southwest.
-- Prickles straight, rarely curved; sepals commonly eglandular on outer surfaces or margins; rare or absent in highest elevations of Rocky Mtns. and outlying peaks and ridges....subsp. woodsii.


Rosa woodsii Lindley subsp. woodsii..
Rosa demareei E. J. Palmer, J. Arnold Arbor. 10:26. 1929. Syn. nov...
Rosa deserta Lunell, Amer. Midl. Naturalist 2:156. 1912....
Rosa fendleri Crépin, Bull. Soc. Roy. Bot. Belgique 15:91. 1876. Rosa woodsii var. fendleri (Crépin) Rydberg in Bessey, Fl. Nebr., Rosales 21:22. 1895....
Rosa fimbriatula Greene, Leafl. Bot. Observ. Crit. 2:135. 1912...
Rosa foliolosa (var.) leiocarpa Torrey in Frémont, Rep. 89. 1843...
Rosa hypoleuca Wooton & Standley, Contr. U.S. Natl. Herb. 16:131. 1913...
Rosa macounii Greene, Pittonia 4:10. 1899....
Rosa maximilani Nees in Wied-Neuwied, Reise N. Amer. 2:434. 1841...
Rosa naiadum Lunell, Amer. Midl. Naturalist 3:139. 1913...
Rosa poetica Lunell, Amer. Midl. Naturalist 3:138. 1913....
Rosa sandbergii Greene, Leafl. Bot. Observ. Crit. 2:137. 1911...
Rosa standleyi Rydberg, N. Amer. Fl. 22:527. 1918...
Rosa subnuda Lunell, Amer. Midl. Naturalist 2:153. 1912....
Rosa terrens Lunell, Amer. Midl. Naturalist 2:151. 1912. Rosa woodsii var. terrens (Lunell) Breitung, Canad. Field-Naturalist 68:82. 1954. Syn. nov....

(10 Sep 2007)  Page(s) 352.  
 
Localized clusters of glabrous populations on the north foot of the San Bernardino mountains represent the variety glabrata within Rosa woodsii subsp. gratissima.

(10 Sep 2007)  Page(s) 351.  
 
Rosa woodsii Lindley subsp. gratissima (Greene) W. H. Lewis & Ertter, stat. nov. Basionym: Rosa gratissima Greene, Fl. Francisc. 1:73. 1891. Rosa pisocarpa var. gratissima (Greene) Jepson...Rosa woodsii var. gratissima (Greene) D. Cole ...

(10 Sep 2007)  Page(s) 346-347.  
 
Rosa woodsii Lindley, Ros. Monogr. 21. 1820....
Key to the subspecies of Rosa woodsii in Central and Western North America:
- Shrubs most frequently tall, 1-3 (-5) m; flowering branches unarmed or with straight or curved prickles; terminal leaflets ovate to elliptic, fewer obovate; flowers 1 or 2 or corymbose and then 3 to 10 (to 15); western North America including the far Southwest.
-- Prickles straight or occasionally somewhat curved, absent or few to dense; flowers few, 3 most common or numerous (to 15); sepals eglandular or infrequently stipitate-glandular; intermontane between Cascade and Rocky Mtns. of British Columbia to southern Great Basin and mountains of Majave Desert, and east to Rocky Mtns.
--- Stems densely branching; prickles stout and often long, straight or curved, sometimes infrastipular and often dense and varying in size intermodally; SW Great Basin, Mojave Desert and adjacent mountains...subsp. gratissima.

(10 Sep 2007)  Page(s) 351.  
 
Rosa woodsii Lindley subsp. manca (Greene) W.H. Lewis & Ertter, comb. et stat. nov. Basionym: Rosa manca Greene, Pittonia 4:11. 1899....It is not the R. manca sensu Erlanson (1934:230) which is described as a hexaploid ecotype species of R. nutkana (= subsp. melina).

(10 Sep 2007)  Page(s) 346.  
 
Rosa woodsii Lindley, Ros. Monogr. 21. 1820....
Key to the subspecies of Rosa woodsii in Central and Western North America:
- Shrubs most frequently short, typically less than 1 m, infrequently to 2 m; fertile branches densely prickly of various sizes, rarely few or unarmed; terminal leaflets obovate, fewer ovate or elliptic; flowers 1 to 3, rarely more; prairies and plains of central North America extending into the Rocky Mtns. and nearby Southwest.
-- Prickles curved, rarely straight; sepals commonly stipitate-glandular on outer surfaces or margins; endemic to high elevations of Rocky Mtns. and outlying peaks and ridges....subsp. manca

(10 Sep 2007)  Page(s) 350.  
 
Rosa woodsii Lindley subsp. arizonica (Rydberg) W. H. Lewis & Ertter, comb. et stat. nov. Basionym: Rosa arizonica Rydberg, N. Amer. Fl. 22:516. 1918...
Rosa adenosepala Wooton & Standley. Contr. US Natl. Herb 16:131. 1913. Syn. nov....
Rosa bakeri Rydberg. Fl. Colorado 191. 1906. nom. illeg., non Rosa bakeri A. Déséglisé...
Rosa granulifera Rydberg, N. Amer. Fl. 22:517. 1918. Syn. nov. Rosa arizonica var. glandulifera (Rydberg) Kerney & Peebles...
Rosa neomexicana Cockerell, Entomol. News 12:38. 1901. Syn. nov. ...