(1960)  Page(s) 114-118.  
 
SINCE 1932 when I first placed pollen on Rosa macounii, one of the three native rose species of Saskatchewan, on pistils of the hardiest of the Hybrid Rugosa, Hansa, I have been concerned with the inheritance of color in roses.....

Soon after my original cross, I put pollen of Harison's Yellow on pistils of R. macounii and got a number of seedlings. These were pollen-fertile in spite of the fact that one parent is tetraploid and the other diploid. One of the seedlings was a salmon-pink bicolor that combined the yellow of the one parent with the pink of the other, although both colors were relatively pale. The remainder were pure white.

That pure white appeared in these seedlings was unexpected, for surely one would have expected all these seedlings to be a salmon-pink. However, apparently, Harison's Yellow is heterozygous for its yellow color, and R. macounii heterozygous for its pink color. These white seedlings, as it chanced, did not inherit yellow from the one parent, or pink from the other and so had to be white....

Later, I placed pollen from Harison's Yellow on pistils of Rosa spinosissima altaica, the pure white rose from the Altai mountains of central Asia, and from this cross I got many new varieties. They ranged from pure white through various tints of pale yellow to the definite yellow of which two were eventually named Hazeldean and Yellow Altai. In these, the yellow is just a mite less deep than in Harison's Yellow. These two roses probably represent the maximum for yellow that can be expected to descend from the stock of yellow genes in Harison's Yellow to any progeny.