(1975)  Page(s) 59-66.  
 
Abstract from Blackwell Synergy website: "Rosa nanothamnus has uncertain affiliations within the genus Rosa. However, evidence is presented that its system of inheritance is of the distinctive and singular type found in the section Caninae .It is considered that this complex system is unlikely to have arisen purely by fortuitous hybridization but is, in fact, highly evolved. Species which share the canina-type system of inheritance are interpreted as a closely related group. Rosa nanothamnus is therefore placed in section Caninae.
Analysis of chromosomes at pachytene in meiosis in Rosa nanothamnus showed that there were 7 bivalents and 14 univalents. It is suggested that the genes which restrict chromosome pairing in this species, and presumably in the Caninae generally, affect synapsis of chromosomes rather than the formation of chiasmata."